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1990-1995 Assistant Professor, UC San Diego, Biology Department.
1995-1999 Associate Professor (tenured), UC San Diego, Biology Department.
1999-2007 Professor, UC San Diego, Cell and Developmental Biology Department.
1998-2003 Member, Mario Negri Sud Research Institute, Italy.
1999-2000 ICREA Visiting scientist, University of Barcelona, Spain.
2007 ICREA Research Professor and Group Leader at the CRG, Barcelona, Spain / Coordinator of the Cell & Developmental Biology Programme, CRG, Barcelona, Spain